Panama's Brilliant Butterflies: Arrival: Oct. 5th Day 1 - Oct. 6th; Las Minas am/canopy Lodge grounds pm Day 2 - Oct. 7th; Cerro Gaital am/el Valle roadsides pm Day 3 - Oct. 8th; Altos del Maria/Finca Macarena Day 4 - Oct. 9th; Canopy Lodge am/canopy Tower pm Day 5 - Oct. 10th; Metropolitan Park am/canopy Tower pm Day 6 - Oct. 11th; Ammo Ponds/Pipeline Road full-day Day 7 - Oct. 12th; Canopy Tower Naturetrek Panama's Brilliant Butterflies October 5-12, 2018 TOTAL SPECIES: 272 new species added: 4 list compiled by: Linda Harrison PAPILIONIDAE SWALLOWTAILS Papilioninae Battus lycidas Swallowtails & Cattlehearts Cramer's Swallowtail Parides childrenae childrenae Green-celled Cattleheart Parides sesostris tarquinius Emerald-patched Cattleheart Heraclides androgeus Androgeus Swallowtail Heraclides thoas Thoas Swallowtail PIERIDAE WHITES & SULPHURS Dismorphilnae Mimic-Whites Dismorphia amphione praxinoe Tiger Mimic-White Dismorphia crisia lubina Crisia Mimic-White Dismorphia theucharila fortunata Clearwinged Mimic-White Enantia jethys Jethys Mimic-White Coliadinae Sulphurs & Yellows Eurema albula White Yellow Eurema x. xantochlora Tropical Yellow Eurema salome jamapa Salome Yellow
Pyrisitia nise Phoebis argante Pierinae Ascia monuste Leptophobia caesia Pieriballia viardi noctipennis Perrhybris pamela fruhstorferi Archonias brassolis approximata Catasticta h. hegemon LYCAENIDAE Theclinae Eumaeus godartii Evenus regalis Brangas getus Theritas mavors Arawacus tograna Ocaria thales Cyanophrys herodotus Thestius epopea Thestius sp. nov. Laothus barajo Lamprospilus collucia Calycopis calus Strymon serapio Tmolus echion echiolus Panthiades bethildis Oenomaeus ortygnus Polyommatinae Cupido comyntas texana RIODINIDAE Euselasilinae Euselasia aurantia Riodinianae Mesosemia carissima Mesosemia asa Leucochimona lagora Mimosa Yellow Apricot Sulphur Whites Great Southern White Bluish White Viardi White Dimorphic White Cattleheart White Frosted Dartwhite HAIRSTREAKS & BLUES Hairstreaks White-tipped Cycadian Regal Hairstreak Bright Brangas Deep-green Hairstreak Togarna Hairstreak Thales Hairstreak Tropical Greenstreak Epopea Hairstreak Thestius Hairstreak (undescribed) Barajo Hairstreak Two-toned Groundstreak Calus Groundstreak Bromeliad Scrub-Hairstreak Red-spotted Hairsteak Zebra-striped Hairstreak Aquamarine Hairstreak Blues Eastern Tailed-Blue METALMARKS Euselasias Bright-centered Euselasia Metalmarks Blue-patched Eyed-Metalmark Deep-blue Eyed-Metalmark Lagora Metalmark
Perophthalma lasus Lasus Metalmark Perophthalma tullis Tullius Metalmark Hyphilaria parthenis Fire-banded Metalmark Eurybia lycisca Blue-winged Eurybia Eurybia patrona persona Great Eurybia Eurybia e. elvina Blind Eurybia Necyria duellona White-dashed Metalmark Ancyluris j. jurgensenii Costa-spotted Metalmark Ancylirus i. inca Inca Metalmark Rhetus arcius Long-tailed Metalmark Ithomeia eulema imitatrix Northern Mimic-Metalmark Brachyglenis dinora Dinora Metalmark Detritivora barnesi Barne's Metalmark Calephelis fulmen Fulmen Calephelis Calephelis iris Iris Calephelis Parcella amarynthina Orange-banded Metalmark Caria rhacotis Honduran Metalmark Caria mantinea lampeto Lampeto Metalmark Baeotis sulphurea Sulphur Metalmark Mesene phareus Cell-barred Metalmark Esthemopsis c. colaxes Colaxes Metalmark Symmachia xypetemultesima Multesima Metalmark Symmachia tricolor Tricolor Metalmark Symmachia xypete Tiger-striped Metalmark Sarota acantus Simple Sarota Sarota myrtea Godman's Sarota Anteros chrysoprasta Elegant Anteros Anteros formosus Black-bellied Anteros Anteros k. kupris Giant Anteros Anteros renaldus indigator Regal Anteros Emesis lucinda White-patched Emesis Emesis aurimna White-spotted Emesis Emesis lacrines Striated Emesis Emesis ocypore aethalia Dark Emesis Emesis cypria Orange-striped Emesis Thisbe irenea Irenia Metalmark
Thisbe ucubis Juditha odites praeclarum Juditha caucana Calospila lucianus Nymphidium ascolia ascolia Theope virgilius Theope basilea Theope phaeo Theope sp. NYMPHALIDAE Danainae Lycorea halia atergatis Tithorea tarricina Aeria eurimedia Mechanitis lysimnia macrinus Napeogenes peridia hemimelaena Napeogenes tolosa amara Ithomia diasia hippocrenis Ithomia heraldica Ithomia iphianassa Oleria paula Oleria rubescens Cereatinia tutia dorilla Callithomia h. hezia Callithomia hezia smalli Dircenna dero euchytma Dircenna klugii Godyris kedema furina Godyris zavaleta zygia Hypoleria lavinia vanilia Greta morgnae oto Heliconiinae Actinote ozomene Dione juno Dryas iulia Philaethria dido panamensis Ucubis Metalmark Odites Metalmark Molpe Metalmark Lucianus Metalmark Creamy Metalmark Blue-based Theope Straight-lined Theope Falcate Theope Theope sp. BRUSHFOOTS Monarchs & Clearwings Tiger Mimic-Queen Cream-spotted Tigerwing Banded Tigerwing Lysimnia Tigerwing Peridia Tigerwing Tolosa Tigerwing Diasia Clearwing Heraldica Clearwing Iphinassa Clearwing Paula's Clearwing Rubescens Clearwing Tutia Clearwing Hezia Clearwing Hezia Clearwing Dero Clearwing Klug's Clearwing Kedema Clearwing Variegated Clearwing Lavinia Clearwing Rusty Clearwing Longwings Lamplight Actinote Juno Longwing Julia Green Longwing
Eueides aliphera Eueides lineata Heliconius cydno chioneus Heliconius doris Heliconius erato petiverana Heliconius hecale melicerta Heliconius sapho Heliconius sara Limenitidnae Adelpha barnesia leucas Adelpha cytherea Adelpha erotia Adelpha i. iphiclus Adelpha melanthe Adelpha salmoneus salmonides Adelpha zina lacina Biblidinae Catonephele mexicana Myscelia cyaniris Myscelia leucocyana Ectima erycinoides Hamadryas feronia Nica flavilla Dynamine agacles core Dynamine postverta Callicore lyca Callicore tolima Cyrestinae Marpesia berania Marpesia chiron Marpesia merops Marpesia petreus Marpesia zerynthia Nymphalinae Colobura dirce Tigridia acesta Fine-lined Longwing Thick-edged Longwing Cydno Longwing Doris Longwing Crimson-patched Longwing Hecale Longwing Sapho Longwing Sara Longwing Sisters Spot-celled Sister Cytherea Sister Erotia Sister Iphiclus Sister Rayed Sister Golden-banded Sister Zina Sister Banners, Crackers & Eighty-eights East-Mexican Banner Whitened Bluewing Pale Bluewing Northern Ectima Variable Cracker Little Banner Pale Sailor Mexican Sailor Six-spotted Eighty-eight Blue-and-orange Eighty-eight Daggerwings Orange Daggerwing Many-banded Daggerwing Spot-banded Daggerwing Ruddy Daggerwing Waiter Daggerwing Beauties & Crescents Dirce Beauty Tiger Beauty
Hypanrtia lethe Anartia fatima Anartia jatrophae Siproeta epaphus Siproeta stelenes Chlosyne narva Chlosyne janais Anthanassa d. drusilla Anthanassa tulcis Castilia eranites Castilia ofella Eresia clio Eresia ithomioides alsina Tegosa anieta anieta Charaxinae Siderone galanthis Memphis arginussa eubaena Memphis philumena indigotica Memphis kingi Archaeoprepona amphimachus Archaeoprepona meander Archaeoprepona demophon Archaeoprepona demophoon Satyrinae Antirrhea philaretes lindigii Morpho cypris Morpho helenor peleides Morpho menelaus amathonte Morpho theseus Brassolis isthmia Caligo illioneus Caligo telamonius menus Opsiphanes quiteria Selenophanes josephus Pierella luna luna Cissia labe Orange Mapwing Banded Peacock White Peacock Rusty-tipped Page Malachite Narva Checkerspot Crimson-patch Checkerspot Orange-patched Crescent Pale-banded Crescent Mimic Crescent White-dotted Crescent Clio Crescent Variable Crescent Black-bordered Crescent Leafwings Red-striped Leafwing Mottled Leafwing Orange-striped Leafwing King's Leafwing White-spotted Prepona Three-toned Prepona One-spotted Prepona Two-spotted Prepona Morphos, Owls & Satyrs Northern Antirrhea Cypris Morpho Common Morpho Menelaus Morpho Stub-tailed Morpho Small-spotted Owl-Butterfly Dusky Owl-Butterfly Pale Owl-Butterfly Scalloped Owl-Butterfly Josephus Owl-Butterfly Moon Satyr Labe Satyr
Cissia pompilia Cissia pseudoconfusa Euptychia westwoodi Hermeuptychia hermes Magneuptychia libye Pareuptychia ocirrhoe Forsterinaria neonumpha Pseudodebis celia Taygetis uzza Yphthimoides renata HESPERIIDAE Eudaminae Tarsoctenus corytus gaudialis Phanus marshalli Hyalothyrus neleus pemphigargyra Ehtneus matho aequatorius Epargyreus exadeus Epargyreus clavicornis Urbanus proteus Urbanus esmeraldus Urbanus dorantes Urbanus teleus Urbanus tanna Urbanus simplicius Urbanus chalco Astraptes talus Astraptes fulgerator Astraptes alardus latia Astraptes chiriquensis Narcosius nazaraeus Narcosius sp. Autochton longipennis Autochton zarex Spathilepia clonius Cogia calchas Telemiades a. antiope Plain Satyr Gold-stained Satyr Westwood's Satyr Hermes Satyr Blue-gray Satyr White Satyr White-dotted Satyr Celia Satyr Uzza Satyr Renata Satyr SKIPPERS Longtails, Flashers, Scarlet-eyes Gaudy Skipper Common Phanus Dimorphic Skipper Giant Entheus Broken Silverdrop Small-spotted Silverdrop Long-tailed Skipper Esmeralda Longtail Dorantes Longtail Teleus Longtail Tanna Longtail Plain Longtail Great White Longtail Green Flasher Two-barred Flasher Frosted Flasher Chiriqui Flasher Nazareus Flasher Narcosius Flasher Spike Banded-Skipper Sharp-banded Skipper Falcate Skipper Mimosa Skipper Plötz's Telemiades
Pyrginae Myscelus amystis hages Myscelus belti Pyrrhopyge evansi Jemadia pseudognetus Celaenorrhinus e. eligius Nisoniades rubescens Bolla cupreiceps Staphylus vulgata Staphylus ascalaphus Gorgythion begga Potamanaxas thoria Ebrietas osyris Eantis thraso Achlyodes pallida Quadrus cerialis Quadrus c. contubernalis Pythonides jovianus amaryllis Milanion marciana Paramimus stigma Spioniades abbreviata Paches loxus enophanes tryxus Pyrgus oileus Heliopetes alana Hesperiinae Turesis complanula Cobalus calvina Cobalus virbius Damas clavus Damas immacula Carystoides l. lebbaeus Zenis jebus Saliana esperi Saliana saladin Saliana salius Spreadwing Skippers Widespread Myscelus Belt's Myscelus Evan's Firetip Dot-collared Firetip Eligius Flat Purplish Tufted-Skipper Copper-headed Sootywing Golden-snouted Sootywing Central American Sootywing Variegated Skipper Thoria Skipper Yellow-patched Bent-Skipper Southern Sicklewing Pale Sicklewing Common Blue-Skipper Striped Blue-Skipper Variable Blue-Skipper Marciana Skipper Stigma Skipper White-tipped Skipper Glorious Blue-Skipper Glassy-winged Skipper Tropical Checkered-Skipper Alana White-Skipper Ruby-eyes & Grass Skippers Complanula Skipper White-rayed Ruby-eye White-edged Ruby-eye Violet-washed Skipper Immaculate Ruby-eye Lebbaeus Ruby-eye Purple-stained Skipper Perching Saliana Violet-tipped Saliana Sullied Saliana
Saliana severus Thracides thrasea Aroma aroma Synapte silius Anthoptus epictetus Lento hermione Lento xanthina Callimormus juventus Callimormus radiola Remella rita Eprius veleda Mnestheus ittona Vehilius stictomenes illduens Vehilius vetula Parphorus decora Papias dictys Niconiades incomptus Vettius lafrenaye pica Vettius diversa Eutychide complana Onophas columbaria Enosis immaculata Wallengrenia otho Pompeius pompeius Anatrytone mella Vacerra egla Vacerra cervara Vacerra caniola Dark Saliana Thrasea Skipper Aroma Skipper Rain-forest Faceted-Skipper Trailside Skipper Hermione Skipper anthina Skipper Juventus Skipper Radiant Skipper Rita's Remella Veleda Skipper Ittona Skipper Pasture Skipper Vetula Skipper Decora Skipper Bottom-spotted Skipper Half-tailed Skipper Two-toned Skipper Diversa Skipper Compliant Skipper Blue-glossed Skipper Immaculata Skipper Southern Broken-Dash Pompeius Skipper Mella Skipper Egla Skipper Cervara Skipper Caniola Skipper