An analysis of the meroplankton assemblages of Station L4 and the development and application of molecular techniques to aid taxonomic resolution.

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University of Plymouth PEARL https://pearl.plymouth.ac.uk 04 University of Plymouth Research Theses 0 Research Theses Main Collection 202 An analysis of the meroplankton assemblages of Station L4 and the development and application of molecular techniques to aid taxonomic resolution. Highfield, James http://hdl.handle.net/0026./7 University of Plymouth All content in PEARL is protected by copyright law. Author manuscripts are made available in accordance with publisher policies. Please cite only the published version using the details provided on the item record or document. In the absence of an open licence (e.g. Creative Commons), permissions for further reuse of content should be sought from the publisher or author.

Eddystone Eddystone Eddystone Abundance Biomass (g) Abundance 2/05/2008 2/05/2008 23/07/2008 Rep Rep Rep PROTOZOA D290 Lucernariopsis campanulata D0560 Virgularia mirabilis D0750 Cerianthus lloydii juv 3 0.0009 2 D0750 Cerianthus lloydii D30 ACTINIARIA D3250 Halcampa chrysanthellum D340 Edwardsia claparedii F0 TURBELLARIA G NEMERTEA 0.0005 2 G620 Cerebratulus HD NEMATODA 9 0.00 8 K20 Loxosoma indet 3 L80 Sagitta 0.00 N SIPUNCULA indet N90 Golfingia elongata (big) 0.766 3 N90 Golfingia elongata 0.000 N09 Golfingia vulgaris (big) N09 Golfingia vulgaris (small) N60 Nephasoma minutum N279 Phascolion strombus (small) N279 Phascolion strombus (large) O5 ECHIURIDA juv POLYCHAETE BITS P200 Pisione remota 0.000 P270 Aphrodita aculeata j P490 Acholoe squamosa P600 Alentia gelatinosa P970 Harmothoe P020 Harmothoe extenuata 6 0.0059 P040 Harmothoe fraserthomsoni 58 P070 Harmothoe impar 2 0.002 Malmgrenia indet Malmgrenia andreanapolis P00 Malmgrenia mcintoshi 0.0432 5 P90 Malmgrenia glabra P200 Malmgrenia arenicola 4 0.00 36 Malmgrenia marphysae Pholoe indet P690 Pholoe inornata 4 Pholoe baltica 3 P80 Sigalion mathildae P870 Sthenelais boa P890 Sthenelais limicola P2000 Phyllodocidae indet 0.0003 Phyllodocidae longipes P2050 Eteone longa agg

P2540 Anaitides lineata P2530 Anaitides groenlandica P2550 Anaitides longipes P2560 Anaitides maculata P2580 Anaitides rosea P2680 Eulalia indet P2790 Eulalia mustela 0.0008 P2820 Eumida indet P2830 Eumida bahusiensis P2850 Eumida sanguinea P2960 Nereiphylla rubiginosa 7 0.009 P3060 Paranaitis kosteriensis P470 Glycera indet P4720 Glycera alba (big) Glycera alba (small) P4730 Glycera celtica # (>5cm) P4760 Glycera lapidum agg 6 0.0888 7 P4790 Glycera rouxii P480 Glycera tridactyla P4870 Glycinde nordmanni P4870 Glycinde nordmanni (small) P490 Goniada P4930 Goniada maculata P500 Goniadella gracilis? 3 0.0029 2 P5080 Ephesiella abyssorum 2 0.00 4 P5320 Hesionidae indet P540 Podarkeopsis capensis 0.000 Gyptis 2 0.008 P5520 Kefersteinia cirrata 3 0.0038 5 Kefersteinia cirrata # 6 0.0025 6 P5680 Ophiodromus flexuosus P5830 Syllidia armata P620 Ancistrosyllis groenlandica 2 0.0042 P6350 Syllidae larvae P6380 Eurysyllis tuberculata 4 0.0009 3 P660 Trypanosyllis coeliaca 4 0.0026 P6620 Trypanosyllis zebra P6660 Typosyllis indet 7 0.0049 P6700 Typosyllis hyalina P6860 Eusyllis blomstrandi P6980 Odontosyllis ctenostoma 7 0.008 P7000 Odontosyllis gibba P700 Pionosyllis 5 0.002 P7280 Syllides benedicti P7440 Exogone hebes P7450 Exogone naidina P7460 Exogone verugera P750 Sphaerosyllis bulbosa 52 0.05 3 P7520 Sphaerosyllis erinaceus P7555 Sphaerosyllis taylori 5 0.0002 P7560 Sphaerosyllis tetralix 3 0.0002 P760 Autolytus 3

P7840 Proceraea P8030 Nereididae juv P8200 Micronereis variegata 0.000 P8260 Neanthes irrorata P8340 Nereis longissima 2 0.007 2 P8670 Nephtys Bit P8670 Nephtys juv Nephtys indet Nephtys # P8675 Nephtys assimilis P870 Nephtys hombergii P870 Nephtys hombergii # P8720 Nephtys kersivalensis P8740 Nephtys incisa P9050 Pareurythoe borealis 8 0.006 P990 Euphrosine foliosa P9330 EUNICIDA Eunice indet P9680 Eunice # 2 0.0522 P9420 Hyalinoecia tubicola P950 Onuphis sp P9680 Eunice sp indet P9720 Eunice pennata 8 P990 Nematonereis unicornis P9830 Marphysa juv P9840 Marphysa bellii P9840 Marphysa bellii # Abyssoninoe sp? Scoletoma indet 2 P9970 Lumbrineriopsis paradoxa 0.0062 P000 Lumbrineris P000 Lumbrineris # 2 P0020 Lumbrineris agastos P0080 Lumbrineris gracilis 8 0.0636 P0080 Lumbrineris gracilis (big) 3 P00 Lumbrineris latreilli P020 Lumbrineris magnidentata P0280 Arabella # Arabella iricolor P0660 Ophryotrocha Schistomeringos P0970 Parougia caeca P040 Protodorvillea kefersteini P50 Schistomeringos rudolphi P50 Scoloplos P520 Scoloplos armiger P430 Orbinia sertulata P550 Paraonidae indet # P580 Aricidea minuta P650 Aricidea catherinae P660 Aricidea cerrutii P790 Levinsenia gracilis

P850 Paradoneis lyra P2080 Apistobranchus P220 Poecilochaetus serpens P2270 Aonides oxycephala 0.0076 P2280 Aonides paucibranchiata 0.0036 P2480 Laonice indet P2500 Laonice bahusiensis 0.0046 P2690 Minuspio cirrifera 8 0.002 6 P2700 Minuspio cf. multibranchiata Polydora P2760 Polydora caeca # agg P3020 Prionospio fallax P30 Pseudopolydora cf. paucibranchiata P320 Pseudopolydora pulchra P3350 Spio decorata P3360 Spio filicornis P3430 Spiophanes bombyx P3440 Spiophanes kroyeri P370 Chaetopteridae juv P3620 Magelona alleni P3630 Magelona filiformis P3640 Magelona minuta Magelona jonstonii P3750 Chaetopterus variopedatus P3980 Caulleriella zetlandica Caulleriella alata Caulleriella indet P4030 Chaetozone setosa Chaetozone christei P4040 Chaetozone gibber 7 P4230 Tharyx # Tharyx indet P3970 Tharyx killariensis 2 P4280 Aphelochaeta # A P430 Monticellina dorsobranchialis P4630 Cossuridae P4790 Diplocirrus glaucus P5300 Capitella P5530 Heteromastus filiformis P5620 Notomastus (big) P5620 Notomastus (small) 0.000 3 P562 Notomastus (size not measured) P5580 Mediomastus fragilis P5690 Peresiella clymenoides P590 Maldanidae indet P6060 Asychis biceps P6230 Clymenura P6300 Euclymene droebachiensis P6330 Euclymene oerstedii P6380 Heteroclymene robusta (Big) P6380 Heteroclymene robusta (Small) P6380 Heteroclymene robusta # 2

P6430 Johnstonia clymenoides P6480 Praxillella affinis P6700 Notoproctus spp? P7330 Asclerocheilus intermedius P7425 Scalibregma celticum P7430 Scalibregma inflatum P800 Polygordius lacteus 5 0.04 27 P8280 Galathowenia oculata Galathowenia indet P8360 Owenia fusiformis P8430 Amphictene auricoma 0.078 P8540 Lagis koreni 0.000 P8860 Melinna palmata P9040 Ampharete juv P900 Ampharete lindstroemi P960 Amphicteis gunneri P9740 Sosane sulcata Terebellidae indet P9830 Trichobranchidae indet (v.small) P9900 Terebellides stroemi P9950 Trichobranchus glacialis P9960 Trichobranchus roseus P2080 Hauchiella tribullata P270 Polycirrus 2 P2390 Streblosoma intestinalis P2030 Lanice conchilega P2050 Neoamphitrite affinis P20760 Pista cristata 0.0039 P20860 Pista lornensis P20900 Terebella # P2440 Thelepus cincinnatus P2500 Sabellidae juv indet P2850 Euchone juv P22040 Jasmineira caudata 3 0.009 2 P22440 Potamethus murrayi P22550 Pseudopotamilla reniformis P22720 Serpulidae P22880 Hydroides norvegica P23090 Serpula vermicularis P24860 Tubificoides amplivasatus P24882 Tubificoides insularis P24890 Tubificoides pseudogaster agg P260 Grania 0.002 6 Q0 PYCNOGONIDA juv Q70 Achelia echinata agg Q450 Callipallene brevirostris 2 Q600 Anoplodactylus juv Q600 Anoplodactylus indet Q620 Anoplodactylus petiolatus Q640 Anoplodactylus virescens R940 COPEPODA 9 0.003 2 R3590 MYODOCOPA(soft) 8

R35680 PODOCOPIDA(hard) 0 0.0027 0 R3020 Lepeophtheirus pectoralis S60 Nebalia bipes S460 Mysidae juv 3 S660 AMPHIPODA indet S700 Apherusa juv S700 Apherusa 4 S70 Apherusa bispinosa S750 Apherusa jurinei S760 Apherusa ovalipes S930 Eusirus longipes 0.005 S2330 Pontocrates altamarinus S2280 Perioculodes longimanus S2400 Synchelidium maculatum S2450 Westwoodilla caecula S250 Pleustidae indet? S2720 Amphilochoides boecki S2730 Amphilochoides serratipes S2790 Amphilochus manudens 2 S2800 Amphilochus neapolitanus S2880 Gitana sarsi 2 0.000 S3060 Peltocoxa brevirostris Leucothoe richiardi S330 Leucothoe incisa S3330 Metopa juv 8 0.0005 S3700 Stenothoe marina S4280 Urothoe brevicornis S4290 Urothoe elegans S4380 Harpinia antennaria S440 Harpinia pectinata S4470 Parametaphoxus fultoni 20 0.007 25 S4480 Metaphoxus pectinatus S4640 Lysianassidae juv 0.000 S5040 Lepidepecreum longicorne S5270 Normanion quadrimanus 2 0.0002 2 S5390 Orchomene nanus 2 S5860 Austrosyrrhoe fimbriatus 2 0.0023 S6070 Argissa hamatipes S640 Stegocephaloides christianiensis S6260 Iphimedia minuta S6280 Iphimedia obesa S6350 Odius carinatus S6590 Liljeborgia kinahani 0.000 S6600 Liljeborgia pallida 0.0048 S6660 Listriella mollis S680 Atylus falcatus S6820 Atylus guttatus S6830 Atylus swammerdamei S6840 Atylus vedlomensis S6890 Dexamine indet S6900 Dexamine spinosa S6960 Guernea coalita 7 0.0022 7

S7070 Ampelisca S7070 Ampelisca juv S7090 Ampelisca armoricana S700 Ampelisca brevicornis S7200 Ampelisca tenuicornis S780 Ampelisca spinipes 6 0.023 S7220 Ampelisca typica S740 Bathyporeia elegans S7460 Bathyporeia pilosa S7470 Bathyporeia sarsi S7480 Bathyporeia tenuipes S7900 Megaluropus agilis S8050 Melitidae indet S880 Ceradocus semiserratus 52 0.0799 S8220 Cheirocratus S8250 Cheirocratus sundevallii S8350 Eriopisa elongata? 4 S8490 Maera juv 35 S8520 Maera loveni 5 0.002 S8530 Maera othonis 4 0.0223 29 S8530 Maera othonis (male) S8790 Amphithoe ramondi S8980 Gammaropsis maculata S900 Gammaropsis sophiae S8950 Isaeidae S9230 Photis longicaudata S940 Ericthonius S9440 Ericthonius punctatus S9720 Aoridae 78 0.022 S9860 Leptocheirus 28 S9880 Leptocheirus hirsutimanus S990 Leptocheirus tricristatus 0.005 6 S9950 Microdeutopus juv S070 Corophium juv S0260 Corophium sextonae S0330 Siphonoecetes striatus S380 Gnathiidae praniza S3220 Gnathia oxyuraea 0.0002 S4070 Cirolana cranchii S0720 Caprella acanthifera S0760 Caprella linearis 2 S0840 Pariambus typicus S0840 Pariambus typicus S0890 Parvipalpus capillaceus S0960 Phtisica marina 0.0002 S00 Pseudoprotella phasma S4840 Janira maculosa S5590 Idotea juv 2 0.000 S6740 Bopyridae indet S920 Pseudoparatanais batei S9080 Leptognathia gracilis 2 S930 Tanaopsis graciloides 36 0.00

S20220 Eudorella truncatula S20550 Nannastacus unguiculatus 7 0.0007 3 S20030 Bodotria scorpioides S2020 Iphinoe sp S2050 Iphinoe trispinosa S20480 Cumella pygmaea 20 0.0008 6 S20550 Nannastacus unguiculatus S20720 Pseudocuma longicornis S20890 Diastylidae juv S20980 Diastylis laevis S20960 Diastylis bradyi S200 Diastylis rugosa DECAPODA larvae indet DECAPODA zooae DECAPODA (Euphausiidae larvae?) DECAPODA megalopa DECAPODA (paguriidae larvae?) DECAPODA (Caridean larvae?) S22080 Palaemon elegans 0.0036 S22390 Athanus nitescens Processa indet S22990 Processa canaliculata S2309 Processa modica S23280 Crangonidae S2330 Crangon crangon S2340 Philoceras bispinosus S23900 Callianassa subterranea S23840 Jaxea nocturna S23970 Upogebia deltaura (big) S23970 Upogebia deltaura (small <cm) S23990 Upogebia stellata S23990 Upogebia stellata (small <cm) S24450 Anapagurus indet S24650 Pagurus bernhardus S24860 Galathea intermedia 0.0208 S24890 Galathea dispersa S25020 Pisidia longicornis 2 0.0268 46 S2540 Ebalia granulosa S25430 Ebalia tuberosa S25750 Inachus juv S25760 Inachus dorsettensis S25850 Macropodia rostrata S26260 Atelecyclus rotundatus S26660 Liocarcinus juv 2 S26690 Liocarcinus depurator S26730 Liocarcinus pusillus S27790 Pinnotheres pisum Iona thoracia GASTROPODA indet GASTROPODA indet 2 W550 Leptochiton asellus 0.0004 W740 Lepidochitona cinerea 2

W2670 Hydrobiidae? Rissoa indet W2840 Rissoa interrupta W3040 Alvania W3060 Alvania abyssicola W3090 Alvania cancellata? W300 Alvania cimicoides? W3600 Hyala vitrea W3640 Ceratia proxima W4350 Caecum glabrum 9 0.0035 8 W4420 Turritella communis W4420 Turritella communis juv Turbonilla acuta W5240 Ondina? W5270 Ondina divisa W5450 Odostomia turrita 0.00 W5770 Megastomia conoidea W620 Epitonium turtonis W650 Eulimidae W650 Eulimidae juv 0.000 W6530 Eulima bilineata 0.000 W6890 Vitreolina philippi juv 2 0.000 W7000 Aporrhais pespelecani W7000 Aporrhais pespelecani juv W7260 Crepidula fornicata juv W7700 Naticidae juv W7750 Polinices montagui W7770 Polinices pulchellus W7730 Euspira catena W8440 Buccinum undatum juv W8890 Hinia reticulata W9580 Acteon tornatilis Opistobranchidae indet W9640 Scaphander lignarius W9690 Cylichna cylindracea W9730 Roxania utriculus W9790 Philine aperta W9790 Philine juv W9790 Philine indet # W030 Retusa indet W070 Retusa truncatula 0.002 W0940 Akera bullata? W0420 Hedylopsis spiculifera 0.000 W2720 Doto 2 W390 Acanthodoris pilosa W4650 Cuthona W620 PELECYPODA juv Nucula indet W680 Nucula nitidosa W670 Nucula hanleyi W690 Nucula nucleus W6490 Mytilus juv

W6500 Mytilus edulis juv W6640 Musculus discors W6830 Modiolula phaseolina juv W7020 Striarca lactea W770 Glycymeris glycymeris juv W7740 Pectinidae juv W750 Limatula subauriculata W7780 Palliolum tigerinum W8050 Aequipecten opercularis W830 Anomiidae juv W8380 Myrtea spinifera W8420 Lucinoma borealis W8420 Lucinoma borealis juv W8500 Thyasira W8520 Thyasira flexuosa W8800 Semierycina nitida 2 W8970 Montacuta substriata? W900 Devonia perrieri W9050 Mysella bidentata W90 Tellimya ferruginosa W9530 Goodallia triangularis W9650 Cardiidae juv W9680 Acanthocardia aculeata W9690 Acanthocardia echinata W9690 Acanthocardia echinata juv W9750 Parvicardium exiguum 2 W9780 Parvicardium scabrum W9870 Laevicardium crassum W9870 Laevicardium crassum juv 0.0005 W9970 Mactra W9980 Mactra stultorum W20020 Spisula juv W20060 Spisula subtruncata W20320 Phaxas pellucidus (big) W20320 Phaxas pellucidus juv Angulus indet W20450 Angulus squalidus W2050 Arcopagia crassa W20570 Fabulina fabula W20630 Moerella pygmaea 0.000 W20850 Gari juv 0 0.0022 W20870 Gari fervensis W20890 Gari costulata 2 0.3775 2 W20900 Gari tellinella W2200 Timoclea ovata W2020 Abra alba W2040 Abra nitida W2050 Abra prismatica Lutraria lutraria juv W200 Lutraria lutraria Ensis ensis juv W20250 Ensis ensis

W20 Solecurtus scopula W250 Azorinus chamasolen W2550 Gouldia minima 0.0526 2 Dosinia juv W2640 Dosinia lupinus juv W2660 Dosinia exoleta W2850 Venerupis senegalensis.0508 W2850 Venerupis senegalensis juv W2890 Chamelea gallina juv W2890 Chamelea gallina W2930 Clausinella fasciata 2 0.5287 W2230 Mysia undata W22390 Corbula gibba W2250 Hiatella arctica juv W23480 Thracia juv W23500 Thracia convexa W2350 Thracia phaseolina W2360 Cochlodesma praetenue ZA30 Phoronis ZB30 ASTEROIDEA ZB40 Astropecten irregularis ZB2040 OPHIUROIDEA juv ZB2350 Ophiothrix fragilis ZB2420 Ophiocomina nigra 0.083 3 ZB2830 Amphiura juv 8 0.0053 0 ZB2860 Amphiura chiajei ZB2880 Amphiura filiformis ZB3000 Amphipholis squamata 9 0.08 6 ZB30 Ophiura juv 3 0.004 ZB330 Ophiura albida ZB350 Ophiura ophiura ZB3520 ECHINOIDA juv 0.000 7 ZB3550 Psammechinus miliaris ZB3880 Echinocyamus pusillus 6 0.054 24 ZB400 Spatangus purpureus juv 2 0.0479 4 ZB400 Spatangus purpureus 0.0233 ZB4070 Echinocardium cordatum juv 8 ZB4070 Echinocardium cordatum ZB4640 Leptopentacta elongata ZB5260 Leptosynapta inhaerens ZB5330 Labidoplax digitata # agg Branchiostoma lanceolatum 2 0.868 ZD40 Ascidiella aspersa Total numbers 650 3.5768 678

Eddystone Eddystone Eddystone Eddystone Eddystone Eddystone Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 23/07/2008 23/07/2008 23/07/2008 23/07/2008 23/07/2008 23/07/2008 Rep Rep 2 Rep 2 Rep 3 Rep 3 Rep 4 0.0039 0.0045 4 3 0.005 0.0229 2 0.0022 5 0.033 0.0056 3 0.0064 38 0.0054 36 0.00 0.00 9 0.008 2 0.000 2 0.86 0.3239 5 0.0023 8 0.004 0.0053 0.09 3 0.0033 0.032 2 0.08 8 0.2986 5 0.009 4 0.0089 0.039 6 0.032 9 0.006 0.005 2 0.0028

0.0027 3 0.009 0.053 0.059 7 0.25 8 0.2036 7 0.0027 0.2476 0.0037 4 0.0066 3 0.009 4 0.004 2 0.007 0.0055 0.000 0.000 0.004 4 0.0009 0.004 0.000 0.000 8 6 0.0006 0.0008 2 0.0035 4 0.0 7 0.0024 2 0.000 4 0.0808 4 0.000 0.0068 9 0.008 2 0.0048 7 0.000 0.0006

0.000 0.000 0.0002 0.000 0.022 0.002 3 0.0609 0.0852 3 0.0038 0.345 3 0.0265 8 0.002 0.0006 0.0033 0.06 2 0.0304 0.0058 2 0.039 5 0.365 0.0025 2 0.0026

0.0086 5 0.04 6 0.035 5 0.0042 0.0007 0.000 5 0.004 0.000 0.000 0.0054 5 0.0045 3 0.005 0.002 0.009 0.003 3 0.0009 0.002 0.007

0.0027 2 0.0002 0.000 3 0.0069 96 0.0343 52 0.023 40 2 0.007 0.007 0.000 0.063 76 0.992 9 0.4563 28 0.000 0.0006 0.000 3 0.00 0.000 2 0.0028 9 0.008 5 0.0043 0.000 2

0.002 0.000 3 0.0005 2 0.0002 0.0008 4 0.007 5 0.007 5 0.0003 0.000 0.000 5 0.0002 2 0.0022 0.0054 0.0025 0.0027 2 0.000 9 0.005 20 0.0002 0.000 6 0.005 0.0038 4 0.002 5 0.009 0.0007 35 0.004 36 0.0037 6

0.022 7 0.09 0 0.0343 2 0 0.004 3 0.0039 35 0.0002 0.0054 2 0.0002 5 0.0006 0.053 2 0.035 2 0.004 6 0.0009 7 0.0033 4 0.0055 0.000 3 0.0023 2 0.0004 0.000 0.000 0.0009 0.000 0.000 0.000 0.000 0.0005 0.000 3 0.000 0.000 9 0.0004 5

0.0005 5 0.0003 5 0.0004 5 0.0006 0.0002 0.004 4 0.0003 7 0.0008 0.000 0.0002 2 0.003 2 0.005 0.0055 0.6644 3 0.002 2.7975 0.0557 0.0098 0.0022.2484 0.000 0.000 0.002 0.005

0.002 0.0009 0.002 2 0.0002 26 0.0065 5 0.000 0.000 0.004 0.0002 0.000 0.000 4 0.006 2

0.003 0.0086 2 6 0.02 9 0.0396 4 0.0028 0.0009 2 0.000 0.006 25.2559 0.003 0.0003 8 0.0034 8 0.0028 0.0032 0.978 2.0336 0.008 2 0.006

.302 0.708 23.503.0306.083 0.0049 2.255 2 2.2469 4 0.0035 0.0002 2 0.0074 0.0025 0.0084 20 0.0047 22 0.007 9 0.0052 3 0.0049 5 0.0055 0.027 0.000 3 0.065 5 0.456 23 0.0959 8 0.027 0.000 3 0.0004 0.3259 0.0 4 0.0068 3 0.0003 0.0253 7.308 5.267 56.7047 497 6.0956 563 4.3624 525

Eddystone Eddystone Eddystone Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 23/07/2008 23/07/2008 23/07/2008 25/05/2008 25/05/2008 03/09/2008 Rep 4 Rep 5 Rep 5 Rep 5 Rep 5 Rep 0.0047 0.0005 0.005 3 0.0023 23 0.2096 28 0.0099 76 0.04 7 0.000 3 0.000.9 2 0.2678 0.0005 8 0.0024 0.0346 8 0.0038 0.0038 0.848 4 0.0 2 0.0677 2 0.0047 0.000

0.000 0.000 0.000 0.0004 0.0006 3 0.2337 4 0.06 2 0.003 6 0.36 0.002 0.00 0.0029 0.000 0.000 0.007 0.000 0.002 0.0047 5 0.0024 0.008 2 0.0006 0.000 2 0.0007 0.089 3 0.0027 4 0.000 3 6 0.000

0.000 0.0006 5 0.054 0.029 2 2 0.0009 0.0379 0.328 5 0.0699 3 0.004 0.027 2 0.009 3 0 0.072 5 0.002

0.005 0.0097 6 0.0034 0.0005 2 0.000 27 0.000 0.08 4 94 0.064 54 6 0.07 78 46 0.059 2 0.000 230 0.6744 658 2 8 0.000 2 3 0.0002 9 6 0.0035 2 0.0078 9 0.0039 2 0.008 36 6

0.0002 9 0.0006 0.0002 3 0.0228 88 0.0576 0.0006 3 0.0074 3 2 8 0.576 20 0.0054 5 0.000 0.0023 0.045 0.076 42 0.0874 0.0003 3 0.0029 0.0026 4 0.002 0.0007

0.0003 2 0.000 0.000 0.0043 4 0.0009 0.000 3 0.005 4 0.00 0.002 0.000 0.094 0.000 2 0.0037 2 0.0006 0.000 0.0002 4 0.0022 0.000 4 0.0005 2 0.0005 3 0.029 0.0006 38 0.0038

0.23 4 0.0428 0.0002 9 2 0.0069 5 2 0.002 0.0204 6 0.0004 0.000 0.032 2 0.0028 0.0009 0.0032 4 0.0008 4 0.009 0.0023 0.003 0.0002 0.00 0.000 6 0.000 2 0.009 8

2 0.0002 5 0.0004 4 0.0005 0.0008 6 0.0005 5 0.000 0.008 0.0494 0.0067 0.0099 0.006 0.0008

0.000 0.002 9 0.009 0.0039 0.36 2.629 5 0.002 0.0797 2 0.3594 3

2 0.0034 8 0.0039 0.0004 0.398 2 0.0044 0.0294 0.000 0.002 0.0099 33 0.003 7 2 0.0373 8 0.0282 3 0.036 0.206 2 2 0.0029 4.437 0.0004 0.000 2 0.005 0.008 3 0.845 0.239 5 0.04 37 2 0.038

0.3098 2.2349 0.492 2 0.048 4 6.64 0.005 2 0.0005 3 0.0054 9 0.0036 0.003 0.000 0.32 44 0.85 0.000 0.003 7 2.443 0.933 3 0.7272 0.0075 5.48 67.979 492

Cawsand Cawsand Cawsand Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 03/09/2008 03/09/2008 03/09/2008 03/09/2008 03/09/2008 03/09/2008 Rep Rep 2 Rep 2 Rep 3 Rep 3 Rep 4 0.005 3 0.0006 0.005 0.0268 33 0.069 26 0.027 55 0.000 7 0.0002 3 0.0006 24 0.26 0.2556 0.059 0.0006 0.0008 0.005

0.2028 4 0.0002 2 0.0004 4 0.0036 5 0.003 5 0.0034 4 0.0038 2 0.0749 0.002 0.006 25 0.005 48 0.0094 43 0.000 6 0.000 7

4 0.0038 2 0.0042 2 0.3027 3 0.399 0.0234 2 0.477 0.000 2 0.000 2 0.0006 2 0.002 0.0008 0.0209 5 0.048 3 2 0.0259 0.007 4 0.0042 0.0048 7 0.085 0.0007 0.000 2 0.0002

0.000 2 0.003 0.002 0.004 2 0.004 3 0.002 0.0009 0.0058 25 0.0034 27 0.005 7 0.0057 0.0052 3 0.0035 7 0.066 6 0.0306 93 0.0457 05 0.037 6 0.048 7 0.068 50 0.0097 79 0.002 2 0.0004 7 0.0027 79 0.0522 2 0.042 43 0.008 0.439 54 0.307 29 0.826 309 0.00 0.0729 0.0042 37 0.086 43 0.036 83 0.0635 4 0.0222 4 0.035 5 0.000 0.0002 3 0.2266 4 0.0388 3 0.0038 4 0.0026 0.098 8 0.000 0.06 0.0009 7 0.0038 27 0.097 33 0.0538 47 0.0269 54

0.0056 34 0.0432 0 0.0068 4 0.0054 7 0.0053 20 0.0039 5 0.0 2 0.0093 2 0.0709 4 0.0092 0.0006 2 0.9504 82 0.5097 08 0.5788 74 0.059 6 0.09 0.0003 2 0.0265 0.0258 3 0.034 0.002 0.0025 0.008 0.000 2 0.0026 2 0.003 6 0.0032 3

0.000 4 0.0045 0.0004 0.0005 0.0022 7 0.003 2 0.000 0.0035 0.000 3 0.0045 6 0.0066 26 0.0098 3 0.000

0.0058 6 0.025 3 0.0072 0.0048 6 0.033 0.02 9 0.004 0.0006 0.00 0.0003 4 0.000 0.000 0.002 0 0.0022 0.000 4 0.000 5 0.0004 0 0.003 3

0.00 2 0.002 0.000 0.0006 5 0.0005 0.0006 6 0.0008 0.0028 0.0055 0.0033 0.000 0.000 0.000

0.002 2.425 2 0.000.5369 5 5.068 0.0237 4 0.0082 5 0.006 2 0.008 0.003 0.000 0.052 7 0.0074 6 0.0226 2

0.0002 2 0.0002 0.0592 30 0.425 4 0.0684 35 0.028 9 0.0357 36 0.077 2 0.0053 0.000 2 0.0004 0.0034 5 0.002 0.0009 2 0.0256 3 0.000 0.4235 2 0.282 0.000 3 0.0007 0.0002 4.07 0.0676 0.0422 27 0.053 3 0.09 6

0.7574 0.3768.5282 0.8783 0.3847 3.2672 0.0055 0.0009 2 0.0009 4 0.008 4 0.076 0.5984 0.004 4 0.0009 0.0006 4 0.0297 0.0069 5 0.3663 5 0.420 2 2.3294 0.8667 0.0378 8.3456 34.7865 049 20.7273 52

Cawsand Cawsand Cawsand Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 03/09/2008 03/09/2008 03/09/2008 4/0/2009 4/0/2009 4/0/2009 Rep 4 Rep 5 Rep 5 Rep Rep Rep 2 3 0.0057 0.002 2 0.002 0.085 0.054 0 0.785 33 0.3036 33 0.0002 4 0.000 0.0294 0.009 0.0074 0.003 0.0034

0.0005 2 0.052 0.0009 0.0003 0.0047 2 0.0064 8 0.0042 2 0.006 4 0.0037 0.0009 8 0.047 2 0.0076 8 0.0033 22 0.0044 32 0.0002 3 0.0002 2 0.000 7 0.0002

0.0045 0.003 0.436 2 0.346 2 0.25 4 0.070 0.0005 0.0002 2 0.03 7 0.037 0.0004 0.0002 0.000 8 0.0052 0.046 25 0.026 20 0.0005 0.005 0.000

0.0003 3 0.000 2 0.008 0.003 0.000 0.0008 0.003 0.0092 3 0.002 3 0.0004 5 0.0007 0.0005 0.0002 0.0037 8 0.0233 9 0.005 6 0.024 72 0.0334 07 0.0355 20 0.028 63 0.002 64 0.032 65 0.0003 2 0.099 2 0.0069 87 0.0505 54 0.34 25 0.205 445 0.2865 293 0.0283 8 0.0023 9 0.0035 6 0.003 0.0045 0.009 0.0009 0.3399 5 0.0238 9 0.0737 9 0.2398 4 0.037 0.0085 2 0.005 7 38 0.0532 0.53 30 0.073 68

0.072 4 0.0053 0 0.037 0.0006 2 0.0004 28 0.0055 35 0.0203 7 0.479 2 0.0209 0.002 0.00 3 0.0566 0.000 0.6325 2.762 87.5732 42 0.0006 0.006 6 0.006 6 2 0.022 2 0.034 3 0.005 0.0003 0.0004 0.000 0.0026 0.0002 2 0.0043

0.0003 0.0023 6 0.00 5 0.00 5 0.002 2 0.008 5 0.006 5 0.0083 0 0.0074 26 0.06 34 0.000

0.00 2 0.0065 0.005 8 0.059 8 0.0079 23 0.0008 0.0009 0.002 0.0005 0.000 0.002 5 0.0004 8 0.00 9 0.0002 8 0.002 7 0.0005 8

0.0002 6 0.0022 4 0.0003 3 0.0003 4 0.0007 2 0.000 0.000 0.0096 0.0047 0.008 0.000

4.633 0.032 0.42 0.0004 2 0.006 2 0.0092 2 0.02 0.000 0.2788 6 0.35 5 0.066 2

0.000 2 0.0323 0.000 0.369 23 0.2 30 0.247 50 0.0637 4 0.082 9 0.0494 4.832 0.247 0.004 2 0.000 4 0.002 0.06 0.536 28 0.079 5 0.037 9

0.746.3937 0.000 0.000 3 0.078.0993 3.9733 0.20 0.0005 4 0.953 0.0032 3 0.002 0.0006 2 0.077 0.333 0.0009 0.0006 20 0.008 6 0.0548 0.068 3 0.678 5 4 0.0038 8 0.508 0.0762 0.042 9.570 783 3.988 339 9.5843

Cawsand Cawsand Cawsand Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 4/0/2009 4/0/2009 4/0/2009 4/0/2009 4/0/2009 4/0/2009 Rep 2 Rep 3 Rep 3 Rep 4 Rep 4 Rep 5 0.052 24 0.036 32 0.000 2 0.000 2 0.248 2 0.073

0.007 0.0022 3 0.005 3 0.023 0.007 0.009 2 0.049 2 0.0047 30 0.0052 6 0.000 7 0.000 2

0.0022 5 0.524 5 0.967 6 0.000 0.000 0.065 0.028 2

0.0005 2 2 0.00 2 0.000 0.056 9 0.035 3 0.0645 66 0.0249 0 0.028 50 0.0089 55 0.0005 2 0.004 0.099 2 0.008 33 0.868 269 0.828 32 0.0054 7 0.0009 5 0.0004 2 0.025 0.005 0.90 0.003 0.0033 7 0.000 0.004 0.006 9 0.638 74 0.869 63

0.048 8 0.0075 25 0.007 0 0.002 9 0.08 0.06 82 0.905 46 0.0034 8 0.0239 3 0.0273 3 0.0099 0.000 0.000 0.002 0.000

0.0009 0 0.004 2 2 0.0009 0.006 4 0.0022 3 0.099 36 0.03 30

2 0.002 0.00 4 0.0043 0 0.0006 0.0005 0.00 2 0.0009 4 0.0005 4 0.0009 5

0.0008 0.0028 3 0.0004 2 0.0002 0.0036 0.006

5 0.008 2 0.000 0.2734 0.0357 2 0.0004 5 0.0036 4 0.002 4 0.000 3 0.0028

0.000 2 0.004 0.293 23 0.0996 39 0.0249 3 0.03 4 0.0007 63.5276 0.0035 3 0.06 7 0.0097 28

0.6 2 0.475 2 0.3922 2 0.002 0.0008 0.83 0.0047 2 0.00 9 0.008 8 0.002 6 0.32 8 0.406 0.0029 0.83.2508 3

Cawsand Cawsand Cawsand Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 4/0/2009 8/07/2008 8/07/2008 8/07/2008 8/07/2008 8/07/2008 Rep 5 Rep Rep Rep 2 Rep 2 Rep 3 0.003 2 0.0273 2 0.0 20 0.2693 6 0.07 5 4.8005 32 0.008 5 0.005 8 0.000 0.5 0.024 0.0398 0.000

0.000 0.0024 2 0.000 5 0.0028 0.033 0.003 2 0.089 0.000 0.0206 2 0.0027 23 0.0033 26 0.000 53 0.0052 27 0.00 6

0.022 0.0007 0.903 4 0.9096 3 0.5262 2 0.000 0.043 0.000 0.0062 5 0.0034 7 0.035 6 0.0007 0.000 0.000

0.0005 2 0.0008 0.000 0.005 0.0047 0.000 0.000 0.0032 7 0.005 8 0.00 0.007 0.003 3 0.0077 0.0055 2 0.066 69 0.0742 89 0.033 93 0.024 63 0.054 38 0.0066 67 0.003 0.006 0.05 9 0.037 32 0.055 35 0.278 373 0.739 96 0.369 9 0.0058 3 0.0004 4 0.0099 32 0.0045 0.0087 4 0.06 7 0.0002 0.000 3 0.0003 2 0.873 0 0.058 4 0.056 3 0.0375 0.0026 5 0.0032 3 0.077 4 0.2 05 0.2079 36 0.0588 63

0.044 2 0.007 3 0.0049 5 2 0.0025 2 0.073 2 0.002 3 0.0009 0 0.059 2 0.037 6 0.336 3 0.0055 0.0003 0.0004.826 59.54 65 0.420 43 0.0093 0.009 0.0077 0.0334 0.0092 4 0.0602 3 0.000 0.000

0.00 0.000 0.000 0.005 4 0.0004 6 0.0033 0.0007 0.02 7 0.0023 2 0.0008 7 0.0056 0.003

0.004 7 0.009 3 0.0097 6 0.0055 22 0.02 3 0.023 5 0.000 2 3 0.0006 0.000 0.000 0.000 0.000 0.0002 5 0.0004 4 0.000 4 0.000 25 0.0024 0.00 2

0.0008 4 0.005 0.000 3 0.000 4 0.0004 0.0002 0.0 0.0005 0.0457 0.0252 0.000

0.003 5 0.0045 3.3624 3 9.3486 0.004 0.2799 0.04 0.2449 2.3054 0.0062 2 0.0052 2 0.0034 0.0064 0.004 9 0.0946 6 0.342 2

0.2453 2 0.207 22 0.0655 27 2 0.0247 5 0.045 0 0.0459 45 0.0426 0.0009 5 0.0669 4 0.0755 2 0.003 0.2057 68 0.5534 36 0.726 7 2 0.0086 8 2 0.566 2 0.022 0.833 0.0204

0.002 0.000 0.9832 2.273 3 0.9495 3 0.03 0.0027 0.0584 0.007 3 0.057 2 0.000 0.0035 3 0.008 2 0.00 8 0.002 2 0.0023 0.8696 0.0933 4 0.3772 9 2 0.0002 2 3 0.0004 0.208 5 0.9783 0.0024

Cawsand Cawsand Cawsand Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 8/07/2008 8/07/2008 8/07/2008 8/07/2008 8/07/2008 8//2008 Rep 3 Rep 4 Rep 4 Rep 5 Rep 5 Rep 0.007 5 0.0405 2 0.006 0.0325 20 0.78 36 0.287 29 0.000 3 0.0002 2 0.787 0.003

0.0002 0.002 0.0022 0.0002 2 0.0002 5 0.032 0.0785 3 0.088 0.000 0.0005 0.0035 57 0.0064 48 0.0056 23 0.0006 5 0.0005 8 0.0005 0

0.000 0.0007 0.0423 0.4372 3 0.382 6 0.5573 8 0.0003 0.000 0.0038 2 0.000 0.000 0.0056 2 0.0057 3 0.026 4 3 0.0009 0.000

0.000 0.0009 0.0007 0.002 2 0.0043 2 0.003 0.0025 3 0.0009 0.0024 5 0.0006 0.0029 2 0.0086 4 0.078 6 0.056 0 0.0423 60 0.0755 32 0.02 59 0.039 47 0.06 56 0.0002 2 0.002 0.00 39 0.073 4 0.042 49 0.62 224 0.792 229 0.2893 35 0.0082 30 0.0045 40 0.083 4 0.03 2 0.027 4 0.0382 5 4 0.002 0.2352 0.0009 2 0.0823 7 0.056 3 0.0002 0.022 2 0.002 9 0.0206 5 0.237 82 0.6588 39 0.538 93

0.0034 7 0.009 8 0.056 4 0.00 2 0.066 0.0005 24 0.0037 6 0.0756 0.025 9 2 0.0624 5 2 0.007 6.674 5.524 75.4379 40 0.008 2 0.0023 3 0.0007 7 0.0742 0.0483 0.05 4 0.0002 2

2 0.0007 2 0.0004 0.000 5 0.000 2 0.008 2 0.0055 8 0.003 0 0.003 23 0.0298

0.033 3 0.0086 2 0.0042 0.005 20 0.0053 4 0.0073 2 0.0002 0.00 7 0.002 0.000 0.0002 0.0002 7 0.0004 4 0.004 7 0.0004 3 0.002 9

0.006 2 0.00 0.002 0.0007 3 0.002 2 0.007 0.000

2.53 0.0903 4.673 5.3033 2 0.0004 3 0.0059 2 0.688 2 0.226 3 0.804 3 0.0005

0.0507 0.022 0.222 2 0.068 20 0.0236 22 0.87 55 0.533 0.0269 2 4 0.0008 0.355 0.364 0.0272 5 0.02 3 0.0449 0.0024 0.0024 5 0.5537 4 0.369 8 0.0539 0.2493

0.003 0.4505 0.0266 2 0.000 0.029 5 2.0525 2 2.0764 3 0.6526 3 0.00 2 3 0.2723 0.028 3 0.0038 9 0.088 5 0.7483 8 0.9763 0.0599 3 0.0068 0.0898 0.000 0.000 0.000 0.5228

Cawsand Cawsand Cawsand Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 8//2008 8//2008 8//2008 8//2008 8//2008 8//2008 Rep Rep 2 Rep 2 Rep 3 Rep 3 Rep 4 0.094 0.439 22 0.007 30 0.246 25 0.000 3 0.000 5 0.000 0.0029 0.00 0.0027 0.0656

0.00 0.000 0.0009 0 0.0033 8 0.0082 6 0.0099 7 0.0302 3 0.08 2 0.03 0.0066 0.0034 0.0046 40 0.0006 48 0.0092 23 0.0004 0 0.0008 9 0.000 5 0.0003

2 0.00 2.5377 3 0.0603 4 0.566 0.086 6 0.379 0.003 0.05 32 0.0387 24 0.08 23 2 0.000

2 0.0006 0.0005 0.0005 0.002 2 0.00 2 0.004 0.008 0.07 8 0.045 3 0.0053 0.064 0 0.047 02 0.0474 89 0.0 38 0.0075 90 0.026 77 0.000 0.006 0.025 36 0.038 4 0.065 37 0.25 384 0.3266 440 0.288 355 0.0005 2 0.0003 28 0.0069 6 0.062 4 0.0328 0.0052 2 2 0.063 0 0.098 7 0.0663 7 0.0093 9 0.023 7 0.003 8 0.80 90 0.2825 66 0.553 77

0.008 6 0.00 9 0.0037 8 0.004 0.000 0.624 3 0.0603 5 0.749 0.0438 0.0057 2 0.004.2322 67.26 05.38 4 0.0009 3 0.002 4 0.058 6 0.053 4 0.246 2 0.000 0.000 0.000 0.000 0.009 0.003

0.00 2 0.0003 0.0006 8 0.006 7 0.0009 4 0.0007 2 0.0006 3 0.009 0.0055 29 0.048 22 0.0093 2 0.000 0.000

6 0.074 0.0007 3 0.0036 8 0.0077 8 0.03 2 2 0.000 0.009 2 0.000 0.0003 4 0.000 0.004 5 0.0009 5 0.0003 2 0.000 6

4 0.0006 2 0.0009 2 0.0008 4 0.0002 0.007 0.0033

0.0003 2 0.003 3.08 5.9446 3 0.748 0.0977 0.0652 2.9256 2.463 0.000 3 0.0022 3 0.047 2 0.94 9 0.076 5 3 0.986

0.0005 0.000 0.0224 0.000 0.04 35 0.424 26 0.2222 22 0.0072 3 0.0085 22 0.083 0.00 2 0.000 7 0.00 0.0009 0.003 3 0.007 0.0038 0.36 0.068 6 0.0034 0 0.0092 0

0.005 0.0282 0.4744 2.35 0.0065 2 0.009.7376 0.9648 0.92 5 0.733 0.989 0.0005 0.0005 2 0.0008 0.007 0.006 0.000 6 0.004 2 0.0003 5 0.000 3 0.000 2 0.000 0.068 5 0.2308 0 0.759 6 0.000 0.05 0.0059 0.049.3277 4.6222 0.025 0.323

Cawsand Cawsand Cawsand Cawsand Cawsand Cawsand Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 8//2008 8//2008 8//2008 4/0/2009 4/0/2009 6/09/2009 Rep 4 Rep 5 Rep 5 Rep 3 Rep 3 Rep 0.003 3 0.009 0.003 3 0.056 8 0.000 7 0.0557 3 2 0.004 2 4

0.038 0.025 0.0026 0.0005 0.0003 2 0.000 0.0008 3 0.002 5 0.042 3 0.027 3 0.0276 2 0.004 30 0.0039 23 0.0032 7 0.0003 3 0.0002 5 0.00 6

0.0072 2 0.008 6 0.604 3 0.4947 0.4938 2 0.000 0.0005 0.000 0.000 0.00 0.000 0.000 2 0.062 2 0.086 6 0.007 4 0.0002 0.000 2

2 0.0006 3 0.0007 6 0.0008 0.0004 2 0.0026 0.000 6 0.0063 6 0.0007 0 0.000 0.0003 4 0.0039 3 0.036 6 0.0422 4 0.0385 84 0.073 8 0.0228 63 0.05 69 0.048 7 0.0007 4 0.096 47 0.036 24 0.047 45 0.230 420 0.24 422 0.665 228 0.0007 6 0.0022 2 0.069 4 0.0306 5 0.092 7 0.000 0.000 2 0.0423 8 0.046 2 0.0777 0.000 0.0034 0.009 7 0.02 5 0.002 20 0.968 95 0.2099 65 0.555 84

0.024 4 0.0045 5 0.02 49 29 0.002 82 0.0304 3 0.0368 5 0.077 5 0.009 2 0.0026.27 9 0.8336 5.7947 8 2 0.000 0.0037 5 0.007 7 2 0.0036 0.000 0.0502 5 0.08 0.0208 0.025 0.004 4

0.0009 0.0003 0.000 0.0006 6 0.00 7 0.00 7 0.0004 2 0.000 0.000 3 0.00 0.04 8 0.07 24 0.078 24 0.000 0.000

0.0026 3 0.005 4 0.06 6 0.039 2 0.0038 2 0.032 3 0.000 0.002 0.0027 46 0.0009 2 0.0004 28 0.003 4 0.0002 4 0.000 5

0.0004 8 0.0034 6 0.000 0.000 5 2 0.004 2 0.053

3.793 2 2.5392 0.0232 5.40 3.8087 3.639 3 9.3942 3.863 0.003 5 0.06 0 0.023 4 0.003 0.96 22 0.0546 9 0.292 39

0.000 0.258 0.72 4 0.827 32 0.855 8 0.0028 4 0.0352 5 0.0494 4 0.035 0.0503 0.95 3 0.259 9 0.00 2.593 0.2 0.0002 0.039 6 0.0062 7 0.0329 23

0.0006 2 0.008 2 0.032.5535 0.0026 6 0.009 2 0.7889 3.8357 0.895 0.0074 9 0.0087 0.004 0.005 2 0.4068 0.0023 0.0042 3 0.0025 33 2 0.000 3 0.0002 2 0.006 0.3303 0.0888 0.5942 2 0.978 0.065 2

Cawsand Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 6/09/2009 04/06/2008 04/06/2008 04/06/2008 04/06/2008 04/06/2008 Rep Rep Rep Rep 2 Rep 2 Rep 3 0.00 30 5.2258 7 34.087 26 0.0335 22 0.0058 6 0.007 4 <.000 7 0.0002 0.0439 0.0004 0.000 0.0408 0.0943 2 0.0035 0.003

0.0005 0.004 0.0008 3 <0.000 0.002

0.0042 0.0855 0.4328 3 0.0994 2 0.3038 3 0.952 2 0.0007 0.0002 0.0032 0.000 0.000

0.0006 3 0.000 2 0.0005 4 2 0.0026 5 0.002 0 0.0006 0.007 2 0.0006 4 0.0047 3 0.063 3 0.043 2 0.0034 2 0.008 0.0333 29 0.02 0 0.006 56 0.00 0.09 0.23 0.0002 0.0004 0.003 3 0.0005 0 0.030 2 0.0007 0.0002 6 0.0002 0.0009 0.939 0.0068 4 0.004 0.0006 3 0.0258 0.97

0.0587 0.008 55 0.2556 5 0.0439 50 0.0045 0.025 0.000 0.000.373 3 0.090 0.086 0.0028 0.026 0.000 0.0004 <.000 58 0.0209 5 0.002

0.006 0.000 0.0003 0.000 0.000 0.009

0.0263 0.0085 0.0003 <0.000 0.0208 0.0003 0.000 0.002 0.0002 0.0005

0.009 0.000 0.0005 0.0049 0.00 0.000 0.000 2 0.004 0.002 2 0.0059 0.004 0.4638.444

0.00 0.0255 3 0.004 0.02 0.058

2 0.057 2 0.05 0.0002 0.084 3 0.0008 0.23 0.575 47 0.042 38 0.0388 24

0.0406.09 0.077 0.046 2 0.007 2 0.0033 4 0.0003 2 0.000 0.0003 0.086 0.0005 35.356 2 0.592 3 306 6.6553 29 7.3043 348

Rame Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 04/06/2008 04/06/2008 04/06/2008 04/06/2008 04/06/2008 6/07/2008 Rep 3 Rep 4 Rep 4 Rep 5 Rep 5 Rep 0.6656 3.3739 34 5.628 43 5.8859 58 0.0029 0.0093 0.0034 40 0.05 56 0.045 25 0.000 7 0.000 6 0.0036

0.003 0.0035 0.095 0.0005 0.000 0.000 2

0.3238 0.0007 0.555 0.036 0.0534 5.2968 4.90 2 0.00 0.000 2 0.0049

0.002 2 0.0005 0.0026 6 5 0.0022 3 0.003 5 0.0085 23 0.048 0.000 3 0.0043 6 0.009 4 0.0 2 0.009 0.003 3 0.0206 4 0.038 53 0.04 60 0.022 44 0.000 2 0.0006 5 0.00 6 0.0004 7 0.000 0.006 6 0.0044 0 0.007 4 0.008 0.00 7 0.0043 2 5 0.004 3

0.23 7 0.0376 54 0.432 3 0.005 0.000 0.000 7 0.0037 39 0.029 50 0.077 3

0.0004 2 0.0002

3 0.07 3

0.00 0.000 0.008 2 0.007 3 0.0006 0.004 0.0022.9776 2.57

0.007 2 4 3.8865 6

0.0056 0.0054 0.0037 0.00 0.0645 48 0.084 32 0.04 5

0.4359 0.004 0.004 0.002 2 0.008 7 0.0054 2 0.000 0.000 8.609 3.9347.253 3 0.248 2.8044 2 0.5547 2.9397 30 8.2485 408.878 233

Rame Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 6/07/2008 6/07/2008 6/07/2008 6/07/2008 6/07/2008 6/07/2008 Rep Rep 2 Rep 2 Rep 3 Rep 3 Rep 4 0.6784 6.2603 5 8.077 2 2.8834 28 0.008 9 0.0074 58 0.04 24 0.000 5 0.000 8 0.000

0.000 0.000.4448 0.0028 3 0.0026 0.000

0.000 0.000 0.0022 0.583 9.242 2 0.4593 5 0.000 0.000

0.002 8 0.006 5 0.002 6 0.0005 4 0.0004 68 0.0555 3 0.0006 0.0004 9 0.0038 5 0.006 2 0.049 4 0.0046 7 0.0333 5 0.033 4 0.0093 42 0.0096 48 0.038 49 0.0008 3 0.00 5 0.0008 2 0.000 9 0.004 5 0.007 2 0.006 2 0.009 2 0.2607 0.006 0.000 0.0007 3 0.00 5 0.009 4

0.0308 0.0272 0.50 38 0.000 3 0.0077 0.0003 0.000 2 0.005 0.0009 3 0.0002 2 0.000 0.0007 2 0.000

0.000 0.0009 0.0009

0.0039 0.0009 0.000 0.000

0.0027 4 0.0034 2 0.0023 2 0.0004 0.9085 0.5623 0.6299 0.006

0.0039 3 0.009 4 0.3437 4.9977 2 0.64 0.00

0.0005 5 0.0007 0.0052 0.0468 4 0.0557 9 0.0686 6

0.027 0.000 2 0.0023 6 0.007 0.0377 0.000 0.0955 0.0064 0.07.8784 2 8.8879 24 23.867 295 6.5097 226

Rame Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 6/07/2008 6/07/2008 6/07/2008 8/09/2008 8/09/2008 8/09/2008 Rep 4 Rep 5 Rep 5 Rep - 5 combined Rep - 5 combined Rep 37.956 8.20 28 0.007 23 0.06 38 0.000 6 0.000

0.008

2 0.003 3.3874 4 0.702 5 0.009 0.000 2

0.008 0.0027 3 0.0005 5 0.0087 5 0.0032 0.0053 2 0.0006 0.0358 5 0.038 27 0.0045 43 0.000 3 0.0005 8 0.002 2 0.004 0.003 0.0008 0.002 0.0005 6

0.7324 0.0203 0.0423 0.055 0.000 0.000

0.0003

0.000 2 4 3

0.00 2 0.0073 0.8092 0.788

0.0029 7.6778 32 0.575 20 8.8776 0.000

0.004 2 0.053 7 0.0209 4

2 0.0033 2.5772 0.704 0.6075 5.09 46 3.545 20 8.8776 96

Rame Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 8/09/2008 8/09/2008 8/09/2008 8/09/2008 8/09/2008 8/09/2008 Rep Rep 2 Rep 2 Rep 3 Rep 3 Rep 4 3.235 28 3.343 24 3.0368 2 0.0094 38 0.028 39 0.3332 24 0.000 9 0.000 7 0.000 0.07 0.000

0.0008 0.000 0.000 0.000

0.000 0.003 2 0.0009 3.879 8.668 3 0.6443 3 0.0005 0.0008 5 0.005 2 0.922 0.9937

0.0004 2 0.0005 3 0.000 0.000 0.000 0.003 4 0.0025 0.000 6 2 2 0.000 2 0.0046 3 0.00 4 0.0265 2 0.0095 8 0.0268 4 0.037 36 0.08 72 0.086 6 2 0.0007 0.00 8 0.0009 6 0.0007 6 2 0.024 4 2 0.003 0.0024 0.0007 4 0.0032 4

0.0348 0.0007 0.000 6 0.006 3 0.000 2 0.002 6 0.0046 2 0.005 0.0206 0.000 4 0.0005 0.004 0 0.0004 39 0.0046 52

0.000

0.006 2 0.009 6 0.000 0.0006 0.0004

0.0004 0.002 0.0008 4 0.0026 2 0.000 0.000 0.0 0.0328 0.0225.363

0.005

0.0005 2 0.002 0.005 0.033 0.098 2 0.000 6 0.008 6

0.0002 2 0.002 9 0.0574 5.4506 2.5787 2 5.9594 0.327 0.282 2 4.2435 2.6964 73 8.0486 262 5.4376 89

Rame Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 8/09/2008 8/09/2008 8/09/2008 06//2008 06//2008 06//2008 Rep 4 Rep 5 Rep 5 Rep Rep Rep 2 0.068 3.0496 44 6.032 26 3.327 26 0.0063 53 0.044 8 0.00 26 6 0.0002 0.000 0.0036 0.0058 0.0027 0.0254

0.0038 0.0007 0.000 0.0003

0.0046 2 0.0023 0.0043.2336 2 0.4889 6.6889 8 0.000 5 0.0006 6 0.002

0.000 3 0.0023 4 0.0079 9 0.009 7 0.0006 0.023 0.000 0.0036 0.0035 7 0.0447 6 0.0332 3 0.0027 93 0.0222 42 0.008 50 0.0022 6 0.0039 0.000 0.0006 9 0.0009 5 0.0005 2 0.0059 0.005 0.468 0.002 2 0.0005 0.000

0.000 0.005 3 0.033 0.000 0.002 5 0.0049 0.0403 0.000 0.000 0.000 0.0059 2 0.0027

0.0006 0.0038 0.000

0 0.0044 7 0.0067 0.0027

0.0003 0.000 0.000 2 0.009 0.000 3 0.0003 0.000 0.076 0.0037 2.62 2.2084 2 0.0022

0.000 23 0.002

0.000 2 0.006 0.0002 0.0028 0.0049 6 0.002 2 0.0006

0.059 3 0.0053 8 0.008 3 3.668 0.000 3.524 2 6.0903.6958 3 2.754 4.6 3.2739 39 7.084 52 7.5257 8

Rame Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 06//2008 06//2008 06//2008 06//2008 06//2008 06//2008 Rep 2 Rep 3 Rep 3 Rep 4 Rep 4 Rep 5 0.037 4.053 43 6.048 23 2.586 27 0.0055 7 0.0023 3 0.008 5 5 0.000 0.000

0.0005 4 0.004 2 0.0063 0.008

.5686 4 0.58 2 0.739 4 2 0.000 4 0.00 3

0.0003 0.000 4 0.0009 2 0.0025 5 0.006 5 0.0006 3 2 0.0029 0.024 3 0.0248 4 0.025 6 0.05 25 0.0056 23 0.0044 9 0.000 0.0005 2 0.0003 5 0.00 8 0.0003 4 0.0007 4 0.000 3 0.000 2 2 0.0003 0.0022 5

0.090 6 0.0037 7 0.45 3 5 0.0036 0.0057 7 0.0307 2 0.000 2

4 0.0006

0.0009 2 0.000 0.000 0.0027 4 0.000

3 0.00 0.003 0.3307

30.222 0.4074 2 0.0026 8 0.0225

2 0.0063 0.0036 9 0.0042

0.048 9 0.0036 4 0.002 7 0.0294 2 4.633 0.998 3 2.644 36.0695 46 7.6605 27 0.7556 67

Rame Rame Rame Rame Rame Rame Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 06//2008 28/0/2008 28/0/2008 28/0/2008 28/0/2008 28/0/2008 Rep 5 Rep Rep Rep 2 Rep 2 Rep 3 5.354 37 5.9889 44 7.097 3 0.002 4 0.290 2 0.0048 2 0.000 6 0.000 2 0.000 2 2 0.0293 0.075

0.0007 0.003 2 0.0073 2 0.0045 4 0.093 2

0.788 5.333 4 0.626 4 0.0022 2 0.002 3 0.0025 3 0.000

0.000 3 0.004 4 3 0.000 0.037 3 0.0024 7 0.0029 9 0.0007 0.0045 3 0.0087 2 0.0076 0.0057 0.005 9 0.04 3 0.0029 30 0.0077 40 0.0092 26 0.008 3 0.005 7 0.0005 4 0.007 8 0.00 8 0.000 3 0.003 6 0.002 6 0.037 6 0.004 5 0.053 4 0.0002 0.004 6 0.0032 4 0.0034 2

3 0.0049 0.0026 3 0.079 0.0024 2 0.00 2 0.024 0.0084 0.0065 0.000 0.000

0.0006 0.0003

0.002 0.0027 9 0.0028

0.0 0.936 3 0.0252 0.0424

0.000 4 0.08 5 58 36.399 2 2.2359 2 5 0.0029 9 0.0075 0.000

0.004 0.00 2 0.00 4 0.077 0.003 2 0.000 2 0.0022 3

0.045 6 0.0045 0.000 4 0.082 0.0446 3.0826 3.4229 5 3.2065 2 9.36 207 46.3265 204 3.5276 74

Rame Rame Rame Rame Rame Hilmars Box Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 28/0/2008 28/0/2008 28/0/2008 28/0/2008 28/0/2008 7/07/2008 Rep 3 Rep 4 Rep 4 Rep 5 Rep 5 Rep 2 3 0.0997 3.739 30 4.903 24 3.5 4 0.0227 32 0.04 3 0.002 0 0.000 4 0.000 3 2.2305 0.0394 2

0.0009 0.0007 2 0.074 2 0.005 6

0.0057 3 0.3308 4 0.62333 4 0.8905 0.0009 3 0.007 9.695 2

0.00 3 0.0005 4 3 0.004 8 0.042 0.0037 0.000 2 0.0004 2 0.020 8 0.037 0.0063 0.0046 3 0.0067 2 0.0006 3 5 0.0079 2 0.005 2 0.00 0.002 5 0.0006 0.000 0.008 4 0.0008 2 0.00 0.04 7 0.0389 0.002 0.4285 0.0065 2 0.00 0.007 6 0.002 2 0.0006 2

2 6 0.000 9 0.0005 0.005 0.0004 3 0.008 5 0.053 2 0.000 4

2 3

0.004 2 0.00 53 5 4 2 4 2 2

2 0.0035 2 3 0.000 8 3.56.96 0.0366

0.0 9 0.0225 0.0009 0.0024 3.456 28 26.677 0.036 3 0.0396 4 0.0328 0.007

2 0.0004 0.003 0.0038 2 0.0066 4 0.009 4 0.005 0.000 3

0.7889 0.004 0.00 3 0.0007 2 0.0009 0.0236 2 0.665 7 7 4.3465 2.642.47 0.593 4.3802 287 40.53053 83 8.2979 43

Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 7/07/2008 7/07/2008 7/07/2008 7/07/2008 7/07/2008 7/07/2008 Rep Rep 2 Rep 2 Rep 3 Rep 3 Rep 4 0.00 3 0.4297 3 3.5836 0.0834 4.3669 0.0037 9 0.007 9 0.0067 6 0.06 5 0.0077 9 0.09 3 0.000 4 0.000 3 2 0.0003 0.0726 2.332 6.034 2.29299 2 0.0035 0.0357 0.0074 2 0.02 0.0045 4

0.084 0.000 0.0005 0.0044 0.004 0.07 2 0.0027 0.0025 4 0.0037 4 0.029 2 2 0.00 0.000 0.000 7 0.008 5 0.0006 2 0.000

0.007 0.0002 0.5376 0.094 0.0253 0.2003 0.0788 2 0.0233 4 0.278 4 2 0.0039 0.000 0.2485 0.374 23 0.79 25 0.009 2 0.036 0.000 0.09 0.0006 0.0007 0.2357 2 0.6664 4 0.5852 3 0.004 2 0.0006 2

0.002 4 0.00 7 0.0039 0.00 0.000 0.958 85 0.0656 62 0.0495 37 3 0.000 0.0003 0.0027 5 0.05 0.000 3 0.0002 0.002 4 0.0005 2 0.003 0.003 0.0008 0.053 5 0.0036 3 0.056 0 0.005 2 0.03 0.002 3 0.002 4 0.002 9 2 0.008 6 0.0032 3 0.0057 0.00 0.0005 2 0.000 4 0.0006 0.0006 0.0006 5 0.3394 5 0.8948 4 0.0072 0.000 0.0003 4 0.002 5 0.0037 9 0.000 0.024 0.0006 0.606.093

2 0.008 0.008 4 0.0047 8 0.042 0.0389 5 0.05 5 0.0254 0.000 0.077 3 0.004 4 0.0082 4 0.0433 0.0226 6 0.0085 2 0.000 2 0.0053 2 0.0038 0.0049 0.053 6 0.0576 2 0.065 0.039 0.0625 0.0055 0.000 0.000 0.000 0.000 0.000 0.0004 2 0.0004 3 0.000 4 0.000

2 0.000 0.0008 0.008 2 0.004 0.000 0.0007 0.000 0.00 0.000 0.00 0.00 0.007 0.0005 0.0003 2 0.00 0.002

0.0303 25 0.0208 7 0.0284 45 0.0064 5 0.008 2 0.0039 0.000 0.0003 0.0002 2 0.0045 0.023 0.0029 8 0.024 0.00 0.00 0.0098 9 0.002 3 0.0022 2 0.008 2 0.0042 8 0.0026 3 0.0008 3 0.005 2 0.0007 0.006 0.000 0.000 2 0.0003 0.0053 8 0.004 0.0023 4 2 0.002 3 0.006 2 0.0004 3 0.0008 0.000

0.0008 0.0005 2 0.002 0.079 0.00 4 0.022 0.008 0.0022 6 0.0009 3 0.007 2 0.0004 0.0022 0.4544 0.603 0.0308 0.5094 0.6902 0.002 0.057 0.0079 2 0.0628 0.000 0.005

0.00 6.5203 3.8492 0.005 0.0038 0.03 3

0.000 0.4 0.08 0.023 3 0.0636 0.000 2.246 2.0298 0.0236 0.099 0.003 8 0.07 0.007 2 3 0.003 0.0099 5 0.03 3 0.000 0.0677 0.0235 6 0.0483 3 0.0248 6 0.004 4 0.668

9.7425 4.9073 0.079 0.0058 0.0058 0.000 2.558 0.0046 0.0033 0.0034 0.0006 8 0.096 3 0.00 8 0.024 6 0.084 2 0.0076 2 0.078 6.9063

Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 7/07/2008 7/07/2008 7/07/2008 7/09/2008 7/09/2008 7/09/2008 Rep 4 Rep 5 Rep 5 Rep Rep Rep 2 0.000 0.007 4 0.004 3.288 2 7.623 0.082 6 0.02 0.0062 5 0.004 2 0.04 0.000 8 0.000 2 0.0002 6 0.0002 0.000 0.000 0.0438 3.6252 3 0.0079 2 3 2 0.003 0.0692 2 0.002 2 0.07 2 0.0042 3

0.000 0.0026 0.0002 6 0.002 0.093 0.067 0.000 4 0.002 2 0.0002 0.005 0.004 6 0.003 3 0.0029 3 2 0.0002 0.000 0.000 0 0.0024 0 0.002 7 2

0.000 0.000 8 0.0046 4 0.076 2 0.7095 2 0.0444 0.93 3 0.08 2 0.2208 2 0.0203 0.0758 0.046 20 0.097 30 0.39 40 0.008 0.000 2 0.000 3 0.8253 0.0843 0.0007 0.0006 3 0.0002

0.000 5 0.00 5 0.0002 5 0.0002 0.000 0.36 06 0.0955 99 0.657 53 0.0002 2 0.000 0.000 3 0.0065 6 0.0038 7 0.0066 2 0.0009 7 0.0026 2 2 0.0008 0.003 2 0.008 2 0.0006 0.003 22 0.0203 4 0.0076 6 3 0.020 3 0.0057 2 0.008 5 0.0022 2 0.0047 0.004 4 0.003 4 0.002 0.005 2 0.0023 0.007 2 0.0006 0.0022 0.0008 0.009 0.0062 0.002 9 0.0089 0.045 4 0.053 6 0.0046 7 0.0029 7 0.005 4 0.00

0.0005 2 0.003 5 0.004 0.0247 0.000 3 0.0007 2 0.00 0.037 2 0.0275 4 0.0374 0 0.0075 7 0.08 0.000 0.0068 0.0024 0.0264 0.0909 0.057 4 0.0365 6 0.0256 0.0005 3.9332 0.0008 0.000 0.003 2 0.0282 0.0002 2 0.0002 0.000 0.000 8 0.0004 20

2 0.000 0.00 2 0.007 3 2 0.0006 0.0005 0.000 2 0.0027 2 0.002 0.000 3 0.0005 4 0.00 0.0026 0.0035 0.0067 2 0.00

0.086 0.3 3 0.027 57 0.0437 39 0.0397 0.0037 0.0007 0.003 0.0006 0.000 2 7 3 8 0.002 2 8 0.00 5 0.0004 68 0.033 2 0.0002 5 0.006 0.0003 0.0007 9 0.003 0.003 0.0007 2 0.000 0.0005 0.009 3 0.0004 0.028 0.002 2 0.0003 0.000 2 0.000

0.000 0.000 0.0002 0.000 0.0009 0.0067 0.000 0.003 2 0.0024 2 0.000 4 0.642 0.0086 0.6797 0.6209 2 0.935 4 2.4853 3 2 0.005 0.024 0.0058 0.023 0.0023 0.0043 0.0008

2 0.57 0.55 2 0.0009 0.000 0.0264 0.0036 0.000

0.000 2.2396 0.024 2 2 0.0344 0.0055 7 0.0094 0.0004 7 0.0627 3 0.0023 0.0002 2 0.000 0.000 0.0542 2 0.0498 2 0.5094 4 0.0254 2 0.063 4 0.05 0.0264 3 0.057 0.009 3 0.0668 2 0.003 2

5.453 2.0778 0.0 0.0067 0.0028 0.000 0.667 0.000 0.0007 0.0005 6 0.0079 3 0.000 0.02 0.0026 3 0.005 24 3.6583 8 0.063 3 438 9.6592 422

Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 7/09/2008 7/09/2008 7/09/2008 7/09/2008 7/09/2008 7/09/2008 Rep 2 Rep 3 Rep 3 Rep 4 Rep 4 Rep 5 0.4072 3 8.3067.742 2.4688 0.002 0.4043 2 0.027 4 0.0037 7 0.0002 5 0.0205 3 0.000 0.4 4.2236 0.047 0.0284 0.043 0.0548 4.8249 3.483.38 0.0004 2 0.000 0.004 0.000 0.0628 0.0333

0.024 0.00 0.0006 0.002 0.000 2 0.000 0.0455 0.000 3 0.0008 0.000 4 0.00 0.0259 3 0.008 0 0.0079 6 0.0053 0.0054 2 0.004 3 0.0048 4 0.007 5 0.003 3 0.0006 4 0.000 0.000

0.0024 4 0.0026 0.0006 9 0.0988 0.0963 0.245 0.82 0.37 2 0.052 0.0548 0.0329 2 0.058 0.2064 33 0.0797 27 0.0 3 5 0.006 0.296 0.0004 0.0004 0.000 0.004 0.3227 2 0.476 2 0.0692 0.000

0.008 5 0.0006 0.000 9 0.000 3 0.0005 6 0.005 4 0.034 7 0.005 48 0.089 74 0.005 0.000 2 0.026 0.004 6 0.0062 7 2 0.003 0.005 7 0.0032 2 0.0059 2 2 0.0024 0.000 3 0.0077 7 0.026 6 0.0086 8 0.0079 2 0.0047 4 0.0084 0.0003 8 0.003 5 0.0005 6.5774 0.00 4 0.002 0.008 3 6 0.0033 0.000 0.0023 0.0025 0.0055 0.6333 2 0.724 2 0.643 0.0579 8 0.0282 3 0.0084 7 0.0038 3 0.00 3 0.0026 3 0.0006 0.003.4768 0.005 0.0225 0.000

0.9802 0.0602 0.024 3 0.0058 0.005 0.000 0.0543 0.0285 3 0.042 6 0.0002 0.006 2 2 0.092 2 4 0.002 0.0045 0.0002 2 0.0398 4 0.0089 0.0028 0.006 4 0.0078 2 0.0536 0.0006 2 0.006 0.002 0.0008 2 0.002 0.0022 2 0.000 0.0002 0.0048 8 0.002 23 0.0035 49 2 0.0006

2 2 0.002 0.000 0.000 0.0005 0.000 2 0.0026 0.000 0.000 0.0044 0.00 0.0006 3 0.000 0.0009 0.0004 0.004

0.022 0.0424 70 0.096 40 0.0438 68 0.0857 2 0.047 0.0026 4 0.0006 0.0002 0.003 0.003 4 0.0007 0.058 4 0.0245 3 0.034 0.0049 4 0.00 0.0057 20 0.093 28 3 0.0004 4 0.007 6 2 0.0006 2 0.0005 6 3 0.00 0.0002 2 0.0009 0.003 0.000 0.000 2

0.000 0.0004 0.0069 2 0.0002 0.005 5 0.0002 2 0.000 2 2.8058.896 2 0.0007 0.025 0.375 0.065 0.0885 0.0722 4 0.0209 0.508

0.0002 0.007 5 2.788 5 3.408 0.0006 2 0.0005 0.0046 0.006 2 0.000

0.005 0.000 0.0026 0.0084 0.004 0.002 2 6 0.004 2 0.023 0.0073 0.635 7 4 0.405 2 0.000 0.004 2

0.00604 2 0.000 2 0.0005 0.000 2 2 0.0007 0.073 8 0.002 5 0.000 3 0.0002 2 0.65 0.0049 4 0.0306 0.0059 2 26.7349 404 9.86994 287.7976 5

Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 7/09/2008 05//2008 05//2008 05//2008 05//2008 05//2008 Rep 5 Rep Rep Rep 2 Rep 2 Rep 3 0.000 2.69 2.274 0.0025 3 0.0032 0.0095 5 0.023 3 0.0002 0.000 5.03 2 0.0078 0.286.293.25 0.0009 0.000 0.794 0.004 2 0.000 2 0.0853 0.0348

0.0206 0.000 2 5 0.0203 3 0.0054 0.002 0.0003 0.000 0.032 0.005 2 0.0382 2 0.002 0.0075 0.000 0.000 6 0.000 4 0.002 0.000 0.006 5 0.005 2 0.0004 3 2 0.000 2 0.0008 2 0.000

0.0308 3 0.04 3 0.004 4 0.074 2 0.934 0.0689 3 0.672 2 0.0284 0.56 0.6 23 0.332 2 0.093 0 9 0.32 0.023 0.0003 0.4375 0.0834 5 0.6897 2 0.0003 2 0.000

0.0023 9 0.0022 4 0.003 6 0.002 3 0.00 7 0.0026 3 0.498 82 0.2222 83 0.72 34 0.0008 0.003 0.0003 0.005 0.0005 0.0036 5 0.0029 6 3 0.0096 5 0.0042 4 0.006 2 0.0233 4 0.0088 4 0.0088 7 0.000 0.0053 0.0082 0.003 0 0.0023 9 0.002 9 0.009 0.000 0.004 2 0.004 0.0006 4 8 0.0033 4 0.0023 2 0.000 3 2 0.65 0.0407 3 0.0699 2 0.0036 4 0.0024 4 0.0046 6 0.0047 4 0.000 0.000 0.62

0.0228 0.000 0.0566 0.0062 2 0.0066 4 0.25 0.0053 0.00 0.0029 0.0678 0.0246 0.09 0.0267 2 0.0236 0.057 0.0029 3 0.0277 2 0.69 2 0.0035 3 0.0005 0.000 0.000 0.000 0.0086 3 0.0007 7 0.00 9

0.0023 0.0002 0.000 6 0.008 4 0.0048 5 3 0.0053 5 0.0095 0.0002 0.005 0.00 3 0.0009 0.0026 3 0.003 7 2 0.008 2 7 0.000 0.0007 0.0007

2 0.073 2 0.0974 63 0.08 52 0.098 6 2 0.0426 2 0.09 0.008 2 0.007 2 0.002 0.005 2 0.00 0.0046 2 0.0028 4 0.0026 0 0.006 4 0.0074 0.0093 8 0.038 87 0.0487 2 2 0.0082 60 0.0324 0.0028 4 0.0009 0.0007 3 0.009 0.0007 0.0007 0.000 4 0.0004 2 9 0.0066 2 0.007 8 0.0004 0.000 3

2 0.002 0.000 0.0004 8 0.0237 3 0.002 22 0.000 2 0.0006 0 0.009 0.0027 3 0.006 4 2.589 4 0.0059 3 6 0.068.355 0.008 0.0443 0.0242 0.0842 0.335 0.0407 0.0077

2 0.0092 2 0.542 0.007 0.000 4 0.0289 0.000 0.02 0.0009 0.04 0.002

0.029 5 2.5597 2 2.548 3 0.0047 2 0.000 0.0004 0.000 0.0074 8 0.0002 0.2426 2 0.00942 2 0.72 4 0.0046 0.002 2 0.005 0.27 0.0005 2 0.089 2 0.0066 3 0.0285

6.6693 0.23 2 0.442 0.073 0.0007 0.0008 0.000 0.00 3 0.0008 3 0.0058 0.004 3 0.0067 0.07 0.000 3 0.0023 7 0.005 9 0.004 0.0049 5.7802 393 8.482 475

Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Hilmars Box Biomass (g) Abundance Biomass (g) Abundance Biomass (g) Abundance 05//2008 05//2008 05//2008 05//2008 05//2008 8/03/2009 Rep 3 Rep 4 Rep 4 Rep 5 Rep 5 Rep 0.0205 2.49 3 0.004 3 0.0065 3 0.0705 0.0083 3 0.0063 8 0.072 0.0002 4 0.000 0.0008 2 0.072 3.476 3.40 0.9406 0.0025 0.0034 0.0005 2 0.0003 0.000 0.0333 3 0.47 2 0.92

0.0823 0.0002 0.00 0.002 0.0002 0.0008 3 0.0033 4 0.0025 3 0.0007 0.0002 0.000 0.005 3 0.0052 3 0.0032 0.006 0.0027 4 0.0076 0 0.0087 3 0.0006 4 0.0003 7 0.008 2 0.000 2 0.000 0.000

0.4383 0.003 5 0.0076 7 0.059 3 0.985 0.555 0.0633 4 0.5429 0.063 0.0369 2 0.0808 0.0293 0.0083 0.025 24 0.244 8 0.0504 2 0.0343 2 0.2453 2 0.0378 0.005 0.98 0.8633 0.0005 4 0.002

0.003 4 0.0007 5 0.004 3 0.005 2 0.0009 3 0.0009 0.2288 99 0.652 73 0.372 37 0.000 0.000 0.0007 0.0024 7 0.007 9 0.002 0.022 5 0.009 2 0.0048 0.0008 2 0.009 2 0.0048 0.0072 4 0.00 6 0.0062 4 0.000 3 0.0029 0.009 0 0.009 6 0.00 5 0.00 0.004 0.002 5 0.006 0.0064 3 0.0032 0.004 0.000 0.000 3 0.003 0.0003 0.0067 0.0005 0.079 0.0055 2 0.2899 0.057 4 0.0063 2 0.639 0.0025 3 0.00 6 0.005 6 0.0023 2 0.007

0.0002 0.094 7 0.0226 5 0.0074 2 2 0.006 0.0073 0.0009 2 0.0069 0.00 0.0378 0.005 2 0.029 0.0579 0.072 0.0048 3 0.0076 2 0.0057 0.0442 0.006 3 0.00 0.000 0.0002 0.0042 56 0.063 37 0.049